A guide to European terrestrial and freshwater species of Enchytraeidae (Oligochaeta)
Keywords:
Clitellata, enchytraeids, key, species identification, taxonomyAbstract
A guide to European terrestrial and freshwater enchytraeid species is presented, designed for the identification of living specimens. Altogether 206 species are included. Recent taxonomic advances regarding new species, revisions, and improved standards of description are integrated. Marine and exotic species are excluded. The illustrated keys are preceded by introductions into the taxonomic traits, the general anatomy of enchytraeids, and the technique of handling living worms during the identification process. Due to persistent taxonomic problems in many groups, a 'sensu lato' approach is adopted in the circumscription of several common species. Bryodrilus ehlersi glandulosus Dózsa-Farkas, 1990 and Enchytraeus christenseni bisetosus Rota & Healy, 1994 are elevated to species rank; the latter receives a new name, Enchytraeus dichaetus, due to homonymy with Enchytraeus bisetosus Levinsen, 1884, now Fridericia bisetosa. Marionina libra Nielsen & Christensen, 1959 is transferred to Bryodrilus. Marionina serbui Botea, 1984 is synonymised with Buchholzia simplex Nielsen & Christensen, 1963. Mesenchytraeus franzi Nurminen, 1977, Euenchytraeus bisetosus Bretscher, 1906 and Cognettia clarae Bauer, 1993 are probably one species. The youngest name is maintained here, pending a systematic revision of the group. The purpose of this key is to facilitate and to stimulate work with enchytraeids.
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References
Abrahamsen, G. (1969): Enchytraeus norvegicus sp. nov.: a new species of Enchytraeidae (Oligochaeta) from Norway. – Nytt Magasin for Zoologi, Oslo 17: 161–164.
Backlund, H. O. (1946): Swedish Enchytraeida I, with anatomical remarks, and with notes on three Lumbricidae. – Kungliga Fysiografiska Sällskapets Handlingar, N.F. 57: 1–24.
Backlund, H. O. (1947): Swedish Enchytraeida II. – Kungliga Fysiografiska Sällskapets Handlingar, N.F. 58: 3–31.
Bauer, R. (1993): Cognettia clarae n. sp. – eine neue Enchytraeiden-Art aus dem österreichischen Fichtenwald (Oligochaeta; Enchytraeidae). – Linzer biologische Beiträge 25: 685–689.
Bauer, R. (1996): On the distribution of enchytraeids and their bioindicative applicability. – Newsletter on Enchytraeidae 5: 11–16.
Bell, A. W. (1936): Three new species of Fridericia (Enchytraeidae) from California. – University of California Publications in Zoology, Berkeley 41: 145–164.
Bell, A. W. (1947): Some new enchytraeids (Oligochaeta) from the old world. – Transactions of the American Microscopical Society 66: 190–202.
Bell, A. W. (1958): The anatomy of the oligochaete Enchytraeus albidus, with a key to the species of the genus Enchytraeus. – American Museum Novitates, New York 1902: 1–13.
Bell, A. W. (1959): Enchytraeus fragmentosus, a new species of naturally fragmenting oligochaete worm. – Science 129: 1278.
Beylich, A. (2005): A method for preservation and mounting of Enchytraeidae (Oligochaeta, Annelida). – Proceedings of the Estonian Academy of Sciences: Biology, Ecology 54: 352–355.
Botea, F. (1984): A new species of Marionina (Oligochaeta–Enchytraeidae) in the caves of Romania: Marionina serbui n. sp. – Travaux de l’Institut de Spéleologie ‘Emile Racovitza’ 23: 3–5. Bouguenec, V. & N. Giani (1987): Deux nouvelles espèces d’Enchytraeus (Oligochaeta, Enchytraeidae) et redescription d’E. bigeminus Niel. & Chr. Remarques sur le genre Enchytraeus. – Annales de Limnologie, Toulouse 23: 9–22.
Bouguenec, V. & N. Giani (1989): Biological studies upon Enchytraeus variatus Bouguenec & Giani 1987 in breeding cultures. – Hydrobiologia 180: 151–165.
Bretscher, K. (1899): Beitrag zur Kenntnis der Oligochaeten-Fauna der Schweiz. – Revue Suisse de Zoologie 3: 369–426.
Bretscher, K. (1900): Mitteilungen über die Oligochaetenfauna der Schweiz. – Revue Suisse de Zoologie 8: 1–44.
Bretscher, K. (1902): Beobachtungen über die Oligochaeten der Schweiz VI. Folge. – Revue Suisse de Zoologie 10: 1–29.
Bretscher, K. (1903): Beobachtungen über die Oligochaeten der Schweiz VII. Folge. – Revue Suisse de Zoologie 11: 1–21.
Bretscher, K. (1906): Über ein neues Enchytraeidengenus. – Zoologischer Anzeiger 29: 672–674.
Buchholz, R. (1862): Beiträge zur Anatomie der Gattung Enchytraeus, nebst Angabe der um Königsberg vorkommenden Formen derselben. – Schriften der königlichen physikalisch-ökonomischen Gesellschaft zu Königsberg 4°. 3. Jahrgang (1863): 93–132.
Cech, G. & K. Dózsa-Farkas (2005): Identification of Fridericia schmelzi sp.n. combining morphological characters and PCR-RFLP analysis. – In: Pop, V. V. & A. A. Pop (eds): Advances in Earthworm Taxonomy II. – Cluj University Press, Cluj-Napoca: 99–118.
Cejka, B. (1912): Die Oligochaeten der Russischen in den Jahren 1900–1903 unternommenen Nordpolarexpedition. II. Über neue Bryodrilus- und Henlea-Arten. – Mémoires de l’Academie Impériale des Sciences de St.-Pétersbourg, 8e série, Cl. Phys.-Math. 29(2): 1–19.
Cejka, B. (1913): Litorea krumbachi n. spec. n. gen. – Ein Beitrag zur Systematik der Enchytraeiden. – Zoologischer Anzeiger 17: 145–151.
Černosvitov, L. (1928a): Die Oligochaetenfauna der Karpathen. – Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere. Jena 55: 1–28.
Černosvitov, L. (1928b): Eine neue, an Regenwürmern schmarotzende Enchyträiden-Art. – Zoologischer Anzeiger 78: 49–62.
Černosvitov, L. (1929): Communication préliminaire sur les Oligochčtes récoltés par M. P. Remy pendant la croisičre arctique effectuée par le ‘Pourquoi-Pas?’ en 1926 sous la direction du Dr. J.-B. Charcot. – Bulletin du Muséum National d’Histoire Naturelle. Paris, 2e s. 1: 144–149.
Černosvitov, L. (1930): Zur Kenntnis der Oligochätenfauna des Balkans. I. Über die Oligochäten aus Bosnien. – Zoologischer Anzeiger 86: 319–333.
Černosvitov, L. (1934a): Note sur la synonymie de quelques espčces d’Enchytraeidés. – Bulletin du Muséum National d’Histoire Naturelle. Paris, 2e s. 4: 373–376.
Černosvitov, L. (1934b): Zur Kenntnis der Enchytraeiden. II. – Zoologischer Anzeiger 105: 295–305.
Černosvitov, L. (1937a): System der Enchytraeiden. – Bulletin de l’Association Russe pour les recherches scientifiques a Prague (section des sciences naturelles et matématiques) 5(34): 263–295.
Černosvitov, L. (1937b): Die Oligochaetenfauna Bulgariens. – Mitteilungen aus dem Königlich Naturwissenschaftlichen Institut. Sofia 10: 69–92.
Černosvitov, L. (1937c): Zur Kenntnis der Enchytraeiden. III. Revision der Friendschen Enchytraeidentypen. – Zoologischer Anzeiger 117: 191–205.
Černosvitov, L. (1937d): Zur Kenntnis der Enchytraeiden. IV. Revision einiger Michaelsenscher Typen.– Zoologischer Anzeiger 118: 273–281.
Černosvitov, L. (1937e): Notes sur les Oligochaeta (Naididées et Enchytraeidées) de l’Argentine. – Anales del Museo Argentino de Ciencias Naturales, Buenos Aires 39: 135–157.
Černosvitov, L. (1945): Oligochaeta from Windermere and the Lake District. – Proceedings of the Zoological Society of London 114: 523–548.
Chalupský, J. (1986): Czechoslovak enchytraeids (Oligochaeta, Enchytraeidae) I. Enchytraeids from an apple orchard by Bavorov in South Bohemia. – Věstnik Československé společnosti zoologické 50: 13–21.
Chalupský, J. (1991): Czechoslovak Enchytraeidae (Oligochaeta). III. Description of a new species of Enchytronia and notes on two species of Marionina. – Acta Societatis Zoologicae Bohemoslovacae 55: 99–113.
Chalupský, J. (1992): Terrestrial Enchytraeidae (Oligochaeta) and Parergodrilidae (Polychaeta) from Sweden, with description of a new enchytraeid species. – Zoologica Scripta 21: 133–150.
Chalupský, J. (1994): Czech Enchytraeidae (Oligochaeta). IV. Description of Enchytronia pratensis sp. n. and a note on Marionina communis. – Acta Societatis Zoologicae Bohemicae 57: 167–172.
Christensen, B. (1961): Studies on cyto–taxonomy and reproduction in the Enchytraeidae. – Hereditas 47: 387–449.
Christensen, B. (1980): Annelida. Animal Cytogenetics 2. (Series Ed. John, B.). – Gebrüder Bornträger, Berlin: 81 pp.
Christensen, B., Hvilsom, B. V. & B. V. Pedersen (1989): On the origin of clonal diversity in parthenogenetic Fridericia striata (Enchytraeidae, Oligochaeta). – Hereditas 110: 89–91.
Christensen, B. & K. Dózsa-Farkas (1999): Invasion of terrestrial enchytraeids into two postglacial tundras: North-Eastern Greenland and the Arctic Archipelago of Canada (Enchytraeidae, Oligochaeta). – Polar Biology 29: 454–466.
Christensen, B. & K. Dózsa-Farkas (1999): The enchytraeid fauna of the Palearctic tundra (Oligochaeta, Enchytraeidae). – The Royal Danish Academy of Sciences and Letters, Biologiske Skrifter 52: 1–37.
Christensen, B., M. Hvilsom & B. V. Pedersen (1992): Genetic variation on coexisting sexual diploid and parthenogenetic triploid forms of Fridericia galba (Enchytraeidae, Oligochaeta) in a heterogeneous environment. – Hereditas 117: 153–162.
Christensen, B., B. V. Pedersen & M. V. Hvilsom (2002): Persisting clone pool differences in sexual/asexual Buchholzia appendiculata (Enchytraeidae, Oligochaeta) as revealed by genetic markers. – Pedobiologia 46: 90–99.
Coates, K. A. (1980): New marine species of Marionina and Enchytraeus (Oligochaeta, Enchytraeidae) from British Columbia. – Canadian Journal of Zoology 58: 1306–1317.
Coates, K. A. (1994): Widespread polyploid forms of Lumbricillus lineatus (Müller) (Enchytraeidae: Oligochaeta): comments on polyploidism in the enchytraeids. – Canadian Journal of Zoology 73: 1727–1734.
Cognetti, L. (1899): Descrizione dell’ Anachaeta camerani, nuova specie della famiglia degli Enchitreidi. – Bollettino dei Musei di Zoologia ed Anatomia comparata della Reale Università di Torino 14(354): 1–4.
Cognetti, L. (1901): Gli Oligocheti della Sardegna. – Bollettino dei Musei di Zoologia ed Anatomia comparata della Reale Università di Torino 16(404): 13–27.
Cognetti, L. (1903a): Enchitreidi del Cadore. – Bollettino dei Musei di Zoologia ed Anatomia comparata della Reale Università di Torino 18(454): 1–3.
Cognetti, L. (1903b): Descrizione di un nuovo Enchitreide (Mesenchytraeus gaudens n. sp.). – Bollettino dei Musei di Zoologia ed Anatomia comparata della Reale Università di Torino 18(453): 1–3.
Collado, R., E. Martínez-Ansemil & N. Giani (1993): Les Oligochètes aquatiques de la Péninsule Ibérique: description de Stylodrilus curvithecus n. sp. (Lumbriculidae) et de Mesenchytraeus lusitanicus n. sp. et redescription de Cognettia hibernica Healy (Enchytraeidae). – Annales de Limnologie, Toulouse 29: 129–138.
Dash, M. C. (1972): New Enchtryaeidae (Oligochaeta) from the Penny Forest soil of British Columbia, Canada. – Annales Zoologici Fennici 9: 15–16.
Delphy, J. (1921): Études sur l’organisation et le développement des Lombriciens limicoles thalassophiles. – Pillu-Roland, Valognes: 137 pp.
Dequal, L. (1912): Descrizione di un nuovo Enchitreide. – Bollettino dei Musei di Zoologia ed Anatomia comparata della Reale Università di Torino 27(652): 1–3.
Dequal, L. (1914): Gli Enchitreidi della Toscana. – Monitore Zoologico Italiano, Firenze 25: 13–24. Didden, W. A. M., H. Born, H. Domm, U. Graefe, M. Heck, J. Kühle, A. Mellin & J. Römbke (1995):
The relative efficiency of wet funnel techniques for the extraction of Enchytraeidae. – Pedobiologia 39: 52–57.
Ditlevsen, A. (1904): Studien an Oligochäten. – Zeitschrift für wissenschaftliche Zoologie, Leipzig 77: 98–480.
Dózsa-Farkas, K. (1970): The description of three new species and some data to the enchytraeid fauna of the Baradla Cave, Hungary. – Opuscula Zoologica Budapest 10: 241–251.
Dózsa-Farkas, K. (1972): Description of three new Fridericia species (Oligochaeta, Enchytraeidae) from Hungarian rendsina soil. – Annales Universitatis Scientiarum Budapestiensis de Rolando Eötvös Nominatae. Budapest 14: 203–209.
Dózsa-Farkas, K. (1973): Ananeosis, a new phenomenon on the life-history of the Enchytraeidae (Oligochaeta). – Opuscula Zoologica Budapest 12: 43–55.
Dózsa-Farkas, K. (1974): A new Fridericia species (Oligochaeta: Enchytraeidae). – Acta Zoologica Academiae Scientiarum Hungaricae 20: 27–32.
Dózsa-Farkas, K. (1986): Eine neue Enchytraeiden Art (Oligochaeta: Enchytraeidae) aus dem Pilis- Gebirge, Ungarn. – Acta Zoologica Hungarica 32: 281–283.
Dózsa-Farkas, K. (1988a): Fridericia berninii sp. n. und weitere Angaben über die Enchytraeiden-Fauna (Oligochaeta) der Balearen. – Acta Zoologica Hungarica 34: 339–344.
Dózsa-Farkas, K. (1988b): Eine neue Enchytronia-Art aus Marokko. – Opuscula Zoologica Budapest 23: 149–151.
Dózsa-Farkas, K. (1989): Fridericia Arten aus Marokko (Oligochaeta: Enchytraeidae). – Acta Zoologica Hungarica 35: 29–39.
Dózsa-Farkas, K. (1990): New enchytraeid species from Sphagnum-bogs in Hungary (Oligochaeta: Enchytraeidae). – Acta Zoologica Hungarica 36: 265–274.
Dózsa-Farkas, K. (1991a): Neue Enchytraeiden-Art aus tieferen Bodenschichten eines Hainbuchen–Eichenwaldes in Ungarn (Oligochaeta, Enchytraeidae). – Acta Zoologica Hungarica 37: 21–25.
Dózsa-Farkas, K. (1991b): Mesenchytraeus kuehnelti sp. n., a new enchytraeid species (Oligochaeta: Enchytraeidae) from a Sphagnum-bog in Hungary. – Opuscula Zoologica Budapest 24: 97–101.
Dózsa-Farkas, K. (1992): List of enchytraeid synonyma. – Newsletter on Enchytraeidae 3: 16–46.
Dózsa-Farkas, K. (1995): Enchytraeus dudichi sp. n., a new fragmenting Enchytraeus species from Iran (Enchytraeidae, Oligochaeta). – Opuscula Zoologica Budapest 27–28: 41–44.
Dózsa-Farkas, K. (1995b): Self-fertilization: an adaptive strategy in widespread enchytraeids. – European Journal of Soil Biology 31: 207–215.
Dózsa-Farkas, K. (1997): Timmodrilus gen. n., a new genus from the family Enchytraeidae (Oligochaeta). – Opuscula Zoologica Budapest 29–30: 49–52.
Dózsa-Farkas, K. (1999): Taxonomical problems in enchytraeids (Oligochaeta) from Spitsbergen. – In: Schmelz, R. M. & K. Sühlo (eds): Newsletter on Enchytraeidae No. 6. Proceedings of the 3rd International Symposium on Enchytraeidae, Osnabrück, Germany. – Universitätsverlag Rasch, Osnabrück, Germany: 21–30.
Dózsa-Farkas, K. (2002a): The enchytraeid fauna (Annelida, Oligochaeta: Enchytraeidae) of the Fertö- Hanság National Park. – In: The fauna of the Fertö-Hanság National Park. – Hungarian Natural History Museum, Budapest: 153–163.
Dózsa-Farkas, K. (2002b): Notes on the genus Oconnorella Rota, 1995. – Natura Jutlandica Occasional Papers 2: 86–90.
Dózsa-Farkas, K. (2005): Fridericia eiseni sp. n., a new enchytraeid species close to Fridercia ratzeli
(Eisen, 1872). – Proceedings of the Estonian Academy of Sciences: Biology, Ecology 54: 279–291.
Dózsa-Farkas, K. (2009): Review of the Fridericia species (Oligochaeta: Enchytraeidae) possessing two spermathecal diverticula and description of a new species. – Journal of Natural History 43: 1043–1065.
Dózsa-Farkas, K. & G. Boros (2005): Achaeta antefolliculata sp.n., a new enchytraeid species (Oligochaeta, Enchytraeidae) from the rock grassland of the Sas-Hegy in Hungary. – Acta Zoologica Academiae Scientiarum Hungaricae 51: 279–285.
Dózsa-Farkas, K. & G. Cech (2006): Description of a new Fridericia species (Oligochaeta: Enchytraeidae) and its molecular comparison with two morphologically similar species by PCR- RFLP. – Zootaxa 1310: 53–68.
Dózsa-Farkas, K., E. Rota, B. Healy & T. Timm (1998): Estonian Enchytraeidae (Oligochaeta) 1. Terrestrial Enchytraeidae from the Võrtsjärv Limnological Station and from Puurmani. – Proceedings of the Estonian Academy of Sciences: Biology, Ecology 47: 235–246.
d’Udekem, J. (1854): Description d’une nouvelle espèce d’Enchytraeus. – Bulletins de l’Académie Royale des Sciences de Belgique 21: 853–865.
Dumnicka, E. (1976): Enchytraeus dominicae sp.n. – a new species of Enchytraeidae (Oligochaeta) from Poland. – Acta Hydrobiologica, Kraków 18: 421–424.
Dumnicka, E. (1977): Enchytraeus polonicus sp. n., a new species of Enchytraeidae (Oligochaeta) from a cave in the Kraków-Czestochowa Upland. – Bulletin de l’Academie Polonaise des Sciences, Série des sciences biologiques Classe II 25: 163–166.
Dumnicka, E. (1979): Remarks on invertebrate fauna of the Cave Provatina (Greece) with a description of a new species of the family Enchytraeidae (Oligochaeta). – Bulletin de l’Academie Polonaise des Sciences, Série des sciences biologiques Classe II 27: 1041–1046.
Dumnicka, E. (1996): Aquatic Oligochaeta and Aphanoneura from the souterrains of Central Europe with descriptions of a new Enchytraeus species. – Mémoires de Biospéologie 23: 167–171.
Dumnicka, E. (2004): A description of Cernosvitoviella tridentina, a new species of Enchytraeidae (Oligochaeta) from the Italian Alps. – Annales de Limnologie, Toulouse 40: 133–137.
Dunger, W. & H. J. Fiedler (eds) (1997): Methoden der Bodenbiologie, 2nd ed. – Gustav Fischer, Jena: 539 pp.
Eisen, G. (1872): Om några arktiska Oligochaeter. – Öfversigt af Kongliga. Vetenskaps-Akademiens Förhandlingar 1: 119–124.
Eisen, G. (1878): Redogörelse för Oligochaeter samlade under de Svenska expeditionerna till Arktiska trakter. – Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar 3: 63–79.
Eisen, G. (1879): On the Oligochaeta collected during the Swedish expeditions to the arctic regions in the years 1870, 1875 and 1876. – Kongliga Svenska Vetenskaps-Akademiens Handlingar 15(7): 10–49.
Erséus, C. (1976): Littoral Oligochaeta (Annelida) from Eyjafjördur, north coast of Iceland. – Zoologica Scripta 5: 5–11.
Erséus, C. & D. Gustafsson (2009): Cryptic speciation in clitellate model organisms. – In: Shain, D. H. (ed.): Annelids in Modern Biology. – Wiley-Blackwell, Hoboken, NJ: 31–46.
Erséus, C. & B. Healy (2001): Oligochaeta. – In: Costello, M. J., C. S. Emblow & R. White (eds): European Register of Marine Species. A checklist of the marine species in Europe and a bibliography of guides to their identification. – Publications Scientifiques du M.N.H.N., Paris: 231–234.
Erséus, C., E. Rota, T. Timm, R. Grimm, B. Healy & S. Lundberg (2005): Riverine and riparian clitellates of three drainages in southern Sweden. – Annales de Limnologie, Toulouse 41: 183–194.
Friend, H. (1899): New British annelids. – The Zoologist, series 4 3: 262–265.
Friend, H. (1911): New British Henleas. – The Zoologist, series 4 15: 464–468.
Friend, H. (1913a): British enchytraeids. V. Species new to science. – Journal of the Royal Microscopical Society, London 1913: 255–271.
Friend, H. (1913b): Some Jersey oligochaets. – The Zoologist, series 4 17: 456–464.
Gadzińska, E. (1974): Cernosvitoviella parviseta sp. n. – a new species of Enchytraeidae (Oligochaeta) from the Polish Tatra Mountains. – Bulletin de l’Académie Polonaise des Sciences, Série des sciences biologiques Classe II 22: 403–406.
Giani, N. (1979): Description de deux nouvelles espèces d’Enchytraeidae des Pyrenées. – Annales de Limnologie, Toulouse 15: 107–112.
Giere, O. (1974): Marionina istriae n.sp., ein mariner Enchytraeide (Oligochaeta) aus dem mediterranen Hygropsammal. – Helgoländer wissenschaftliche Meeresuntersuchungen 26: 359–369.
Goodrich, E. S. (1897): Notes on Oligochaetes with the description of a new species. – The Quarterly Journal of Microscopical Science 39: 51–69.
Graefe, U. (1980): Systematische Untersuchungen an der Gattung Achaeta (Enchytraeidae, Oligochaeta). 1. Beschreibung von Achaeta brevivasa sp. n. und Achaeta camerani (Cognetti). – Mitteilungen aus dem hamburgischen zoologischen Museum und Institut 77: 35–39.
Graefe, U. (1989): Systematische Untersuchungen an der Gattung Achaeta (Enchytraeidae, Clitellata). 2. Beschreibung von vier neuen Arten. – Mitteilungen aus dem hamburgischen zoologischen Museum und Institut 86: 127–131.
Graefe, U. (2002): On the body architecture of the genus Achaeta (Enchytraeidae, Clitellata). – Natura Jutlandica Occasional Papers 2: 106.
Graefe, U. (2007): Achaeta diddeni sp. nov. (Enchytraeidae, Clitellata) from northern German lowlands. – Folia Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis, Biologia 110: 35–40.
Graefe, U., K. Dózsa-Farkas & B. Christensen (2005): Achaeta unibulba sp. n., a widespread European species (Oligochaeta, Enchytraeidae). – Proceedings of the Estonian Academy of Sciences: Biology, Ecology 54: 271–278.
Healy, B. (1975): A description of five new species of Enchytraeidae (Oligochaeta) from Ireland. – Zoological Journal of the Linnean Society 56: 315–326.
Healy, B. (1979a): Three new species of Enchytraeidae (Oligochaeta) from Ireland. – Zoological Journal of the Linnean Society 67: 87–95.
Healy, B. (1979b): Records of Enchytraeidae (Oligochaeta) in Ireland. – Journal of Life Sciences of the Royal Dublin Society 1: 39–70.
Healy, B. (1979c): Review of the genus Guaranidrilus (Oligochaeta, Enchytraeidae) with the description of two new species. – Bulletin of the British Museum of Natural History (Zoology). London 37: 7–15.
Healy, B. (1980): Records of Enchytraeidae (Oligochaeta) from Western France and the Pyrénées. – Bulletin du Muséum National d’Histoire Naturelle, Paris 4° sér. 2: 421–443.
Heck, M. & J. Römbke (1991): Two new species of Achaeta (Enchytraeidae, Oligochaeta) from meadow and pasture soils of Germany. – Zoologica Scripta 20: 215–220.
Henle, F. G. J. (1837): Ueber Enchytraeus, eine neue Anneliden–Gattung. – Müllers Archiv für Anatomie, Physiologie und Wissenschaftliche Medizin, Berlin 1837: 74–90.
Hoffmeister, W. (1843): Beitrag zur Kenntnis deutscher Landanneliden. – Archiv für Naturgeschichte 1843: 183–198.
International Commission on Zoological Nomenclature (1999): International Code of Zoological Nomenclature, 4th ed. – The International Trust for Zoological Nomenclature, London: 305 pp.
ISO International Standard Organization (2008): Soil quality – Sampling of soil invertebrates – Part 3: Sampling and soil extraction of enchytraeids (ISO 23611–3: 2008(E)). – Guideline prepared by Technical Committee ISO/TC 190, Soil quality, Subcommittee SC 4, Biological methods: 1–12.
Issel, R. (1904): Due nuove Fridericia. – Atti della Società Ligustica di Scienze Naturali e Geografiche, Genova 15: 31–39.
Issel, R. (1905a): Oligocheti inferiori della fauna italica – Zoologische Jahrbücher. Abteilung für Anatomie und Ontogenie der Tiere, Jena 22: 451–476.
Issel, R. (1905b): Materiali per una fauna dell’Arcipelago Toscano, Isola d’Elba. III. Enchitreidi dell’Isola d’Elba. – Annali del Museo di Storia Naturale di Genova (3) 2: 5–8.
Issel, R. (1905c): Un enchitreide ad ampolla spermatecale unica – Atti della Società dei Naturalisti e Matematici, Modena 7: 77–79.
Jansson, B. (1961): Michaelsena glandulifera n. sp., a new enchytraeid fom the interstitial fauna of sandy beaches. – Arkiv för Zoologi, Stockholm 13: 81–91.
Kasprzak, K. (1972a): Achaeta seminalis sp. n., a new species of Enchytraeidae (Oligochaeta) from Poland and notes on other species of the genus Achaeta Vejdovský, 1877. – Bulletin de l’Académie Polonaise des Sciences, Série des sciences biologiques Classe II 20: 187–191.
Kasprzak, K. (1972b): A new species of Enchytraeidae (Oligochaeta) found in the National Park of Great Poland. – Bulletin de l’Académie Polonaise des Sciences, Série des sciences biologiques Classe II 20: 563–566.
Kasprzak, K. (1979): Skąposzczety (Oligochaeta) Pienin. I. Wazonkowce (Enchytraeidae). – Fragmenta faunistica, Warszawa 24: 7–56.
Kasprzak, K. (1986): Skąposzczety wodne i glebowe, II. Rodzina: Wazonkowce (Enchytraeidae). – Polska Akademia Nauk Instytut Zoologii, Warszawa: 366 pp.
Klungland, T. & G. Abrahamsen (1981): Cernosvitoviella ampullax sp.n. (Oligochaeta, Enchytraeidae) from Hardangervidda, S Norway. – Zoologica Scripta 10: 251–253.
Knöllner, F. H. (1935a): Ökologische und systematische Untersuchungen über litorale und marine Oligochäten der Kieler Bucht. – Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere. Jena 66: 425–512.
Knöllner, F. H. (1935b): Die Oligochaeten des Küstengrundwassers. – Schriften des Naturwissenschaftlichen Vereins von Schleswig–Holstein 21: 135–139.
Kobetičová, K. & J. Schlaghamerský (2003): On the efficiency of three schemes of enchytraeid wet funnel extraction. – In: Didden, W. & P. van Vliet (eds): Newsletter on Enchytraeidae No. 8. Proceedings of the 5th International Symposium on Enchytraeidae, Wageningen, The Netherlands, 12–14 April, 2002. – Department of Soil Quality, Wageningen University, Wageningen, The Netherlands: 25–32.
Košel, V. (1975): A new species of Fridericia (Oligochaeta, Enchytraeidae) with asymmetric spermatheca from Slovakia (Western Carpathians). – Biológia, Bratislava 30: 333–336.
Koßmagk-Stephan, K. J. (1983): Marine Oligochaeta from a sandy beach of the Island Sylt (North Sea) with description of four new enchytraeid species. – Mikrofauna des Meeresbodens 89: 1–28.
Koßmagk-Stephan, K. J. (1985): Systematik, Faunistik und Lebenszyklus mariner Oligochaeta der Nord- und Ostsee. Dr.rer.nat. Dissertation, University Göttingen: 306 pp.
Kowalewski, M. (1916): Marionina tatrensis M. Kowalewski (1914) 1916, nowy przedstawiciel rodziny Enchytraeidae. – Rozprawy Akademii Umiejętności: Wydział mat.-przyrodnicy – Kraków, Ser. III 16B: 1–8.
Lasserre, P. (1968): Présence du genre Achaeta Vejdovský, 1877 (Oligochaeta, Enchytraeidae) dans les plages sableuses marines. Description de Achaeta littoralis n. sp. – Bulletin du Muséum National d’Histoire Naturelle, Paris, 2° sér. 39: 979–983.
Leidy, J. (1882): On Enchytraeus, Distichopus and their parasites. – Proceedings of the Academy of Natural Sciences of Philadelphia 1882: 145–148.
Levinsen, G. M. R. (1884): Systematisk-geografisk Oversigt over de nordiske Annulata, Gephyrea, Chaetognathi og Balanoglossi. II – Videnskabelige Meddelelser fra Naturhistoriske Forening i København 1883: 92–350.
Martínez-Ansemil, E. & R. Collado (1996): Two new species of freshwater Oligochaeta from the North- west Iberian Peninsula: Krenedrilus realis sp. nov. (Tubificidae) and Cernosvitoviella bulboducta sp. nov. (Enchytraeidae). – The Journal of Zoology, London 240: 363–370.
Michaelsen, W. (1886): Über Enchytraeus möbii Mich. und andere Enchytraeiden. – Lipsius & Tischer, Kiel: 52 pp.
Michaelsen, W. (1887): Enchytraeiden-Studien. – Archiv für Mikroskopische Anatomie 30: 366–378. Michaelsen, W. (1888): Beiträge zur Kenntnis der deutschen Enchyträiden-fauna. – Archiv für
Mikroskopische Anatomie 31: 483–498.
Michaelsen, W. (1889a): Oligochaeten des Naturhistorischen Museums in Hamburg. I. – Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 6: 1–17.
Michaelsen, W. (1889b): Synopsis der Enchytraeiden. – Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg 11: 1–59.
Möller, F. (1971): Systematische Untersuchungen an terricolen Enchytraeiden einiger Grünlandstandorte im Bezirk Potsdam. – Mitteilungen aus dem Zoologischen Museum in Berlin 47: 131–167.
Möller, F. (1974): Achaeta segmentata n. sp. (Enchytraeidae, Oligochaeta), eine neue Enchytraeidenart aus Lössboden. – Mitteilungen aus dem Zoologischen Museum in Berlin 50: 179–181.
Möller, F. (1976): Achaeta hallensis nom. nov. pro Achaeta segmentata Möller, 1974. – Mitteilungen aus dem Zoologischen Museum in Berlin 52: 175.
Moszyński, A. (1933): Description d’une nouvelle espèce d’Oligochètes Fridericia stephensoni n. sp. – Bulletin International de l’Académie Polonaise des Sciences et des Lettres. Classe des Sciences Mathematiques et Naturelles Série B: Sciences Naturelles (2) Année 1932: 363–367.
Moszyński, A. (1938): Marionina paxi, ein neuer Borstenwurm von den Haberwiesen. – Beiträge zur Biologie des Glatzer Schneeberges 4: 343–345.
Müller, O. F. (1774): Vermium Terrestrium et Fluviatilium, seu Animalium Infusoriorum, Helminthicorum et Testaceorum, non Marinorum, Succincta Historia. Volume 1, Part 2. – Heineck & Faber, Havniae et Lipsiae: 72 pp.
Nakamura, Y. (1993): A new fragmenting enchytraeid species, Enchytraeus japonensis from a cropped Kuroboku soil in Fukushima, Northern Japan (enchytraeids in Japan 5). – Edaphologia 50: 37–39.
Nielsen, C. O. & B. Christensen (1959): The Enchytraeidae. Critical revision and taxonomy of European species (Studies on Enchytraeidae VII). – Natura Jutlandica 8–9: 1–160.
Nielsen, C. O. & B. Christensen (1961): The Enchytraeidae. Critical revision and taxonomy of European species. Supplement 1. – Natura Jutlandica 10: 1–23.
Nielsen, C. O. & B. Christensen (1963): The Enchytraeidae. Critical revision and taxonomy of European species. Supplement 2. – Natura Jutlandica 10: 1–19.
Nurminen, M. (1964): Lumbricillus fennicus sp. n. and some other Enchytraeidae (Oligochaeta) from Finland. – Annales Zoologici Fennici 1: 48–51.
Nurminen, M. (1965): Enchytraeids (Oligochaeta) from northern Norway and western Lappland. – Annales Zoologici Fennici 2: 11–15.
Nurminen, M. (1970a): Records of Enchytraeidae (Oligochaeta) from the west coast of Greenland. – Annales Zoologici Fennici 7: 199–209.
Nurminen, M. (1970b): Four new enchytraeids (Oligochaeta) from southern Finland. – Annales Zoologici Fennici 7: 378–381.
Nurminen, M. (1977): Enchytraeidae (Oligochaeta) from the Grossglockner region of the Australian Alps. – Annales Zoologici Fennici 14: 224–227.
O’Connor, F. B. (1955): Extraction of enchytraeid worms from a coniferous forest soil. – Nature 175: 815–816.
O’Connor, F. B. (1962): The extraction of Enchytraeidae from soil. – In: Murphy, P. W. (ed.): Progress in Soil Zoology. – Butterworths, London: 279–285.
O’Connor, F. B. (1963): Marionina cambrensis sp. nov.: a new enchytraeid worm (Oligochaeta) from North Wales. – The Annals and Magazine of Natural History, including Zoology, Botany and Geology. London 6: 761–766.
Ørsted, A. S. (1844): De regionibus marinis. Elementa topographiae historic-naturalis freti Oeresund. – Inaugural Dissertation, Havniae: 90 pp.
Panchenko, I. (2006): Comparison of two methods for extraction of enchytraeids. – Entomological Review 86 Suppl. 1: S104–S106.
Parapar, J., E. Martínez-Ansemil, C. Caramelo, R. Collado & R. Schmelz (2009): Polychaetes and oligochaetes associated with intertidal rocky shores in a semi-enclosed industrial and urban embayment, with the description of two new species. – Helgoland Marine Research 63: 293–308.
Pfeffer, G. (1890): Die niedere Thierwelt des antarktischen Ufergebietes. – In: Neumayer, G. (ed.): Die internationale Polarforschung 1882–1883. Die Deutschen Expeditionen und ihre Ergebnisse. Band II. Beschreibende Naturwissenschaften in einzelnen Abhandlungen. Hamburg: 455–574.
Piper, S. R., S. F. MacLean & B. Christensen (1982): Enchytraeidae (Oligochaeta) from taiga and tundra habitats of northeastern U.S.S.R. – Canadian Journal of Zoology 60: 2594–2609.
Popčenko, V. (1988): Vodnye maloščcetinkovye červi (Oligochaeta limicola) Severa Evropy (Aquatic Oligochaeta of Northern Europe). – Nauka, Leningrad: 287 pp.
Ratzel, F. (1869): Beiträge zur Anatomie von Enchytraeus vermicularis Henle. – Zeitschrift für wissenschaftliche Zoologie, Leipzig 18: 99–108.
Rodriguez, P. & N. Giani (1986): Description de trois espèces nouvelles d’Oligochètes aquatiques du Pays Basque (Espagne). – Hydrobiologia 139: 269–276.
Römbke, J. (1989): Zur Biologie eines Buchenwaldbodens 12. Die Enchytraeen. – carolinea 47: 55–92.
Rota, E. (1994): Enchytraeidae (Oligochaeta) of western Anatolia: taxonomy and faunistics. – Bollettino di Zoologia 61: 241–260.
Rota, E. (1995): Italian Enchytraeidae (Oligochaeta). I. – Bollettino di Zoologia 62: 183–231.
Rota, E. (2001): Oversized enchytraeids (Annelida, Clitellata): a comparative study, with a revised description of Lumbricillus maximus (Michaelsen). – Organisms, Diversity & Evolution 1: 225–238.
Rota, E. & R. O. Brinkhurst (2000): Mesenchytraeus antaeus, a new giant enchytraeid (Annelida, Clitellata) from the temperate rainforest of British Columbia, Canada, with a revised diagnosis of the genus Mesenchytraeus. – The Journal of Zoology, London 252: 27–40.
Rota, E. & B. Healy (1994): The enchytraeid fauna of North Africa – Hydrobiologia 278: 53–66.
Rota, E. & B. Healy (1999): A taxonomic study of some Swedish Enchytraeidae (Oligochaeta), with descriptions of four new species and notes on the genus Fridericia. – Journal of Natural History 33: 29–64.
Rota, E., B. Healy & C. Erséus (1998): Biogeography and taxonomy of terrestrial Enchytraeidae (Oligochaeta) in Northern Sweden, with comparative remarks on the genus Henlea. – Zoologischer Anzeiger 237: 155–169.
Rota, E., L. Matamoros & C. Erséus (2008): In search of Marionina (Clitellata, Enchytraeidae): A taxonomic history of the genus and re-description of the type species Pachydrilus georgianus Michaelsen, 1888. – Italian Journal of Zoology 75: 1–20.
Schirmacher, A., H. Schmidt & W. Westheide (1998): RAPD-PCR investigations on sibling species of terrestrial Enchytraeus (Annelida: Oligochaeta). – Biochemical Systematics and Ecology 26: 35–44.
Schlaghamerský, J. (1998): The enchytraeids of a beech forest on a basalt-limestone gradient. – In: Pižl, V. & K. Tajovský (eds): Soil zoological problems in central Europe. – České Budějovice: 179–189.
Schlaghamerský, J. (2007): Fridericia brunensis sp. n. (Clitellata: Enchytraeidae) – a new European enchytraeid species similar to F. monochaeta Rota, 1995. – Folia Facultatis scientiarum naturalium Universitatis Masarykianae Brunensis, Biologia 110: 53–65.
Schlaghamerský, J. & V. Pižl (2009): Enchytraeids and earthworms (Annelida: Clitellata: Enchytraeidae, Lumbricidae) of parks in the city of Brno, Czech Republic. – Soil Organisms 81: 11–39.
Schmelz, R. M. (1996): Species separation and identification in the Enchytraeidae (Oligochaeta, Annelida): combining morphology with general protein data. – Hydrobiologia 334: 31–36.
Schmelz, R. M. (1998): Description of Fridericia montafonensis sp. n. (Enchytraeidae, Oligochaeta) from an Austrian meadow. – Mitteilungen aus dem hamburgischen zoologischen Museum und Institut 95: 79–88.
Schmelz, R. M. (1999): On the combined use of morphological and protein patterns in Enchytraeidae species level taxonomy: Example Fridericia discifera Healy, 1975. – Pedobiologia 43: 497–506.
Schmelz, R. M. (2002): Records and taxonomy of Fridericia species found in the Mols area. – Natura Jutlandica Occasional Papers 2: 75–85.
Schmelz, R. M. (2003): Taxonomy of Fridericia (Oligochaeta, Enchytraeidae). Revision of species with morphological and biochemical methods. – Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (Neue Folge) 38: 415 pp. + 73 Figs.
Schmelz, R. M. & R. Collado (1999): Enchytraeus luxuriosus sp. nov., a new terrestrial oligochaete species (Enchytraeidae, Clitellata, Annelida). – carolinea 57: 93–100.
Schmelz, R. M. & R. Collado (2005): Fridericia larix sp. nov. (Enchytraeidae, Oligochaeta) from Irish soils. – Organisms, Diversity & Evolution 5: 85–88.
Schmelz, R. M. & R. Collado (2007): Revision of Hemienchytraeus stephensoni (Cognetti, 1927) (Enchytraeidae, Oligochaeta, Annelida). – Folia Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis, Biologia 110: 67–85.
Schmelz, R. M. & R. M. Collado (2008): A type-based redescription of Pachydrilus georgianus Michaelsen, 1888, the type species of Marionina Michaelsen, 1890, with comments on Christensenia Dózsa-Farkas & Convey, 1997 (Enchytraeidae, Oligochaetae, Annelida). – Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (Neue Folge) 44: 7–22.
Schmelz, R. M. & J. Römbke (2005): Three new species of Hemienchytraeus (Enchytraeidae, Oligochaeta) from Amazonian forest soil. – Journal of Natural History 39: 2967–2986.
Schmelz, R. M., R. Collado & K. Dózsa-Farkas (1999a): New record of Fridericia pyrenaica Giani, 1979, with complements to the original description. – Bulletin de la Société d’Histoire Naturelle, Toulouse 135: 71–75.
Schmelz, R. M., R. Collado & M. Oestreich (1999b): The Enchytraeus buchholzi/christenseni species complex – outline of a problem and first steps towards a solution. – In: Schmelz, R. M. & K. Sühlo (eds): Newsletter on Enchytraeidae No. 6. Proceedings of the 3rd International Symposium on Enchytraeidae, Osnabrück, Germany. – Universitätsverlag Rasch, Osnabrück, Germany: 31–39.
Schmelz, R. M., R. Collado & M. J. I. Briones (2008): Notes on Marionina filiformis (Enchytraeidae, Oligochaeta, Annelida). – Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (Neue Folge) 44: 23–35.
Schmelz, R. M., R. M. Collado de la Peña & J. Römbke (2009): Benefits from ecological study methods to taxonomy of enchytraeids in southern Mata Atlântica. – Pesquisa Agropecuaria Brasileira, Brasilia 44: 861–867.
Schmelz, R. M., N. Arslan, R. Bauer, W. Didden, K. Dózsa-Farkas, U. Graefe, I. Panchenko, A. Pokarzhevski, J. Römbke, J. Schlaghamerský, L. Sobczyk, Z. Somogyi, V. Standen, A. Thompson, J. Ventins & T. Timm (2005): Estonian Enchytraeidae (Oligochaeta) 2. Results of a faunistic workshop held in May 2004. – Proceedings of the Estonian Academy of Sciences: Biology, Ecology 54: 255–270.
Sesma, V. & K. Dózsa-Farkas (1993): Description of seven new species of Enchytraeidae (Oligochaeta) from Spain. – Acta Zoologica Hungarica 39: 249–265.
Shurova, N. M. (1978): The intertidal oligochaetes from the eastern coast of Kamchatka. – In: Litoral Beringova morya i yugo-vostochnoj Kamchatki (Littoral of the Bering Sea and the southeastern Kamchatka). – Nauka, Moscow: 98–105.
Southern, R. (1907): Oligochaeta of Lambay. – The Irish Naturalist’s Journal, Dublin 16: 68–82. Southern, R. (1909): Contributions towards a monograph of the British and Irish Oligochaeta. –
Proceedings of the Royal Irish Academy 27, B(8): 119–182.
Springett, J. A. (1969): A new species of Cernosvitoviella (Enchytraeidae) and records of three species new to the British Isles. – Pedobiologia 9: 459–461.
Springett, J. A. (1971): The Enchytraeidae (Oligochaeta) of South Western Australia: The genus
Fridericia Michaelsen 1889. – Journal of the Royal Society of Western Australia, Perth 54: 17–20. Stephenson, J. (1911): On some littoral Oligochaeta of the Clyde. – Transactions of the Royal Society of Edinburgh 48: 31–65.
Stephenson, J. (1922): The Oligochaeta of the Oxford University Spitsbergen expedition. – Proceedings of the Zoological Society of London 74: 1109–1138.
Stephenson, J. (1926): A description of two Scottish enchytraeids (Oligochaeta). – The Annals and Magazine of Natural History, including Zoology, Botany and Geology. London, series 9 18: 86–91.
Stephenson, J. (1930): On some African Oligochaeta. – Archivio Zoologico Italiano. Torino 14: 485–511.
Stirrup, H. H. (1913): A descriptive study of an oligochaete worm of the family Enchytraeidae; with an appendix on certain commensal protozoa. – Proceedings of the Zoological Society of London 87: 300–321.
Timm, T. (1999): A guide to the Estonian Annelida. – Estonian Academy Publishers, Tartu-Tallinn: 208 pp.
Timm, T. (2005): On the distribution and taxonomic limits of Lumbricillus pagenstecheri (Oligochaeta, Enchytraeidae). – Proceedings of the Estonian Academy of Sciences: Biology, Ecology 54: 292–301.
Timm, T. (2007): Estonian Enchytraeidae 3. Freshwater Enchytraeidae. – Folia Facultatis scientiarum naturalium Universitatis Masarykianae Brunensis, Biologia 110: 87–92.
Timm, T. (2009): A guide to the freshwater Oligochaeta and Polychaeta of Northern and Central Europe. – Lauterbornia 66: 1–235.
Timm, T. & V. Popčenko (1978): The aquatic Oligochaeta from lakes of the Murmansk Region. – Gidrobiologicheskiye issledovaniya (Hydrobiological Researches, Institute of Zoology and Botany of the Estonian Academy of Sciences) 7: 79–111.
Timm, T., C. Erséus & S. Lundberg (1996): New and unusual records of freshwater Oligochaeta from the Scandinavian peninsula. – Nordic Journal of Freshwater Research 72: 15–29.
Ude, H. (1892): Ein neues Enchytraeiden-Genus. – Zoologischer Anzeiger 401: 344–345.
Vejdovský, F. (1878): Zur Anatomie und Systematik der Enchytraeiden. – Sitzungsbericht der Königlich Böhmischen Gesellschaft der Wissenschaften 1877: 294–304.
Vejdovský, F. (1879a): Beiträge zur vergleichenden Morphologie der Anneliden. I. Monographie der Enchytraeiden. – Verlag von Friedrich Tempsky, Prag: 61 pp.
Vejdovský, F. (1879b): Vorläufige Mittheilungen über die fortgesetzten Oligochaetenstudien. – Zoologischer Anzeiger 2: 183–185.
Vejdovský, F. (1884): System und Morphologie der Oligochaeten. – Prag: 172 pp.
von Bülow, T. (1957): Systematisch-autökologische Studien an eulitoralen Oligochaeten der Kimbrischen Halbinsel. – Kieler Meeresforschungen 13: 96–116.
Wachs, B. (1968): Die häufigsten hämoglobinführenden Oligochaeten der mitteleuropäischen Binnengewässer. – Hydrobiologia 30: 225–247.
Welch, P. S. (1917): The Enchytraeidae (Oligochaeta) of the Woods Hole region, Mass. – Transactions of the American Microscopical Society 36: 119–138.
Westheide, W. & U. Graefe (1992): Two new terrestrial Enchytraeus species (Oligochaeta, Annelida). – Journal of Natural History 26: 479–488.
Wilcke, D. E. (1957): Über die verwandtschaftlichen Beziehungen zwischen Enchytraeidenarten aus der Untergattung Enchytraeoides. – Zoologischer Anzeiger 158: 66–71.
Wilcke, D. E. (1967): Oligochaeta. – In: Brohmer, P, P. Ehrmann, G. Ulmer & H. Schiemenz (eds): Die Tierwelt Mitteleuropas, 1. Band, Lief. 7a. Quelle & Meyer, Leipzig: 162 pp.
Xie, Z. & E. Rota (2001): Four new terrestrial species of Marionina (Clitellata, Enchytraeidae) from China and re-examination of M. hoffbaueri Möller. – Journal of Natural History 35: 1417–1431.
Xie, Z. C., Y. L. Liang & H. Z. Wang (2000): A taxonomic study of Bryodrilus (Enchytraeidae, Oligochaeta) from Changbaishan Mountain, China. – Species Diversity 5: 93–101.
Zalesskaya, N. T. (1982): Enchytraeid fauna (Oligochaeta, Enchytraeidae) of the Moscow District. – In: Anonymous (ed.): Soil Invertebrates of the Moscow District. – Nauka, Moscow: 119–132.
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